Wills played in the early days of rock music in Cambridge, from c. 1961 in the Vikings, then a succession of local bands: the Sundowners, Soul Committee, Bullitt (with David Gilmour on guitars and William "Willie" Wilson on drums) and Cochise before joining Frampton's Camel.
Wills joined the Rock band Joker's Wild in 1966, (with David Gilmour on guitars and vocals), replacing Tony Sainty, until they broke up in 1968. He played bass on Peter Frampton's first three albums before parting from Frampton in 1975.
He became the bassist with Roxy Music in 1976, before leaving the band and joining The Small Faces in 1977, during their reunion period. He left the Small Faces and appeared on David Gilmour's eponymous album in 1978, with William Wilson on drums. The next year, Wills became a member of rock band Foreigner and remained with them for fourteen years.
After leaving Foreigner in 1992,he joined Bad Company and stayed with them until Boz Burrell rejoined the band. In July 1999 he filled in for Lynyrd Skynyrd bassist Leon Wilkeson for live shows when Wilkeson briefly took ill. On 24 April 2001, he appeared at The Steve Marriott Memorial Concert, as part of a backing band with Bobby Tench, Zak Starkey and Rabbit Bundrick.
Wills was reunited with The Small Faces drummer Kenney Jones in the Jones Gang during 2006 and appeared with The RD Crusadersfor The Teenage Cancer Trust at The London International Music Show on 15 June 2008.
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